Ray Romano

Manny (voice)

John Leguizamo

Sid (voice)

Denis Leary

Diego (voice)

Queen Latifah

Ellie (voice)

Simon Pegg

Buck (voice)

Seann William Scott

Crash (voice)

Josh Peck

Eddie (voice)

Chris Wedge

Scrat (voice)

Karen Disher

Scratte (voice)

Kristen Wiig

Pudgy Beaver Mom (voice)

Eunice Cho

Madison (Diatryma Girl) (voice)

Harrison Fahn

Glypto Boy (voice)